compiled by Wolfgang A. Nässig
A treatise on the Saturniidae (and Brahmaeidae, Eupterotidae,
Sphingidae etc.) of the island of Borneo (East Malaysia and Indonesia)
was published by J.D. Holloway in 1987:
|Jeremy D. Holloway (1987): The Moths
of Borneo, Part 3: Superfamily Bombycoidea: families Lasiocampidae, Eupterotidae,
Bombycidae, Brahmaeidae, Saturniidae, Sphingidae.
Kuala Lumpur (Southdene), 199 pp., 163 genitalia figs., 20 col. pls.
Availability & prices:
The following species of Saturniidae are known from
arrangement according to the order used in the more recent Sumatran,
which shares a lot of species with Borneo, and Philippine
Saturniidae faunas. Additions or changes in comparison to Holloway’s list
of 1987 are listed as footnotes following the
list. The systematics and nomenclature applied here are therefore not
identical with the original treatment by Holloway in 1987! — See
also the disclaimer below. — References
see at the end.
Holloway (1987) listed 22 species of Saturniidae for the island of Borneo;
here we show 23 species based on the present knowledge.
Superfamily Bombycoidea Latreille, 1802
Family Saturniidae Boisduval,  1834
Subfamily Saturniinae Boisduval,  1834
Tribe Attacini Blanchard, 1840
Genus Attacus Linnaeus, 1767
Attacus atlas (Linnaeus, 1758)
Genus Archaeoattacus Watson, 1914
Archaeoattacus staudingeri (Rothschild,
Genus Samia Hübner, 
Samia (insularis) tetrica (Rebel, 1924)
Tribe Saturniini Boisduval, 
Genus Actias Leach, 1815
Actias maenas (Doubleday, 1847) (see
Genus Cricula Walker, 1855
Actias selene (Hübner, 1806) (see
Cricula elaezia Jordan, 1909
Genus Antheraea Hübner,
Cricula bornea Watson, 1913 (see
Cricula trifenestrata (Helfer, 1837) (see
Subgenus Antheraeopsis Wood-Mason, 1886
Antheraea (Antheraeopsis) youngi Watson,
Hübner,  1816
Antheraea (Antheraea) larissa (Westwood,
Genus Loepa Moore, 1859
Antheraea (Antheraea) platessa Rothschild, 1903
(nec jana (Stoll, 1782)) (see
Antheraea (Antheraea) gschwandneri Niepelt, 1918
Antheraea (Antheraea) brunei Allen & Holloway,
Antheraea (Antheraea) alleni Holloway, 1987 (see
Antheraea (Antheraea) moultoni Watson, 1927 (see
Antheraea (Antheraea) rosieri (Toxopeus, 1940)
Antheraea (Antheraea) helferi borneensis Moore,
Antheraea (Antheraea) diehli Lemaire, 1979
Antheraea (Antheraea) (pernyi) roylii korintjiana
Bouvier, 1928 (see
Antheraea (Antheraea) lampei Nässig &
Holloway, 1989 (see
Loepa megacore Jordan, 1911
Genus Lemaireia Nässig & Holloway, 1987
Loepa sikkima javanica Mell, 1938 (see
Lemaireia loepoides (Butler, 1880)
The taxonomic changes proposed here within this web page are not proposed
as valid taxonomic acts in accordance with the Code; they are only suggestions
to promote the ongoing discussion among the specialists.
References see below!
was listed by Holloway as Samia tetrica. The Samia taxa of
Sundaland (except the second Peninsular Malaysian species?) all belong
to the insularis complex. The interpretation which taxon is to be
treated as species or subspecies remains to be assessed; a revision of
the whole genus is in preparation by R.S. Peigler and S. Naumann. — Back.
status of Bornean (and other Sundanian) Actias maenas populations
requires further research; within the Sumatra
fauna the Sundaland population was called A. maenas diana Maassen,
. — Back.
status of Bornean (and other Sundanian) Actias selene populations
requires further research; within the Sumatra
fauna the Sundaland population was called A. selene seitzi Kalis,
1934. As was recently shown by U. & L.H. Paukstadt (1999a), the populations
of the selene-group from Java and the other parts of Sundaland are
not identical. The valid subspecific name for the Bornean population of
selene may be vandenberghi; but further research remains necessary.
of Cricula bornea is still unclear. — Back.
status of the Bornean population of Cricula trifenestrata as well
as the differences with C. bornea still require further research.
called this species A. assamensis Helfer, 1837. However, the true
(Ao.) assamensis is restricted to the Himalayan area (and maybe mountains
in northern Indochina/South China?) (studies are in preparation, see under
the website www.saturniidae-mundi.de).
In the Sumatra treatment of 1996, this taxon
Watson, 1915 (applied to the Sumatran population) was interpreted as a
subspecies of castanea Jordan, 1910. It was then raised to full
species status by Nässig & Treadaway (July 1998, “The
Saturniidae (Lepidoptera) of the Philippines”). See also U. Paukstadt
et al. (August 1998), who published some additional information on the
Sumatran fauna of Antheraea. — Back.
when Holloway’s Borneo publication was published, the status and identity
of “jana” sensu auctorum was not yet resolved. This was only done
by Nässig (1992). Holloway’s “A. jana” in fact is A. platessa.
The true A. jana is a much smaller species evidently endemic to
Java and Bali. — L.H. & U. Paukstadt (1999) described the larva of
the Bornean A. platessa. It is quite similar to all other species
of the frithi-subgroup, but shows distinctive dichromatic dorsal
scoli on meso- and metathorax in last instar. Regrettably it was not stated
whether the dorsal scoli on A8 are separated in last instar (as illustrated
for earlier instars) or fused. — Back.
called this Bornean population A. celebensis Watson, 1915. This
name celebensis, however, should be restricted to the Sulawesian
population. Calling this Bornean species gschwandneri here in the
present web page follows the concept applied in the Sumatran
web page. However, there are other concepts, see the footnote
in the Sumatran web page. The situation within the species of the frithi-subgroup
(how many real biological species are included in this group, and how can
we differentiate them?) is still far from being solved, see also the following
footnotes. — The species A. ulrichbroschi U. & L.H. Paukstadt,
1999, described from Peninsular Malaysia, may possibly also live on Borneo,
in case that it is a distinct species; compare U. & L.H. Paukstadt
(1999b). — Back.
following footnote. — Back.
and A. moultoni are two more taxa which also belong to the frithi-complex.
Although these populations appear to be easily recognizeable externally,
we still lack a safe confirmation of the species status on a reliable and
well-done methodological base. — Back.
Moore, 1859 was described from India. The status of the Sundanian populations
requires further research. — Back.
publication by Holloway was published in 1987, the identity of these two
taxa was not yet solved; Holloway dealt with both species under the name
of “A. korintjiana Bouvier, 1928” only. In a later paper (Nässig
& Holloway 1989) we separated these two taxa. See also the remarks
in the footnote of the Sumatran publication.
was described first by Moore in 1859, not in his “1858–1859”  publication,
see also the remarks about these publications in the footnote of the Sumatran
publication. The situation within the pernyi-group still has
some problems. — Back.
was called L. sikkima Moore, 1865 by Holloway. However, there are
some differences between the nominotypical northern populations in the
Himalaya area and the Sundanian ones. — Back.
was inadvertedly described within the publication by Holloway (1987) in
advance of the so-intended publication, because the print of the publication
by Nässig & Holloway in Heterocera Sumatrana became delayed into
early 1988. — Back.
Nässig, W. A. (1992):
(Antheraea) platessa Rothschild 1903: The correct name for Antheraea
jana auctorum, nec Stoll 1782 (Lepidoptera: Saturniidae). — Nachr.
entomol. Ver. Apollo, Frankfurt/Main, N.F. 13 (2a): 157–163.
Nässig, W.A., &
Holloway, J.D. (1988) On the systematic position of “Syntherata”
loepoides Butler and its allies (Lep., Saturniidae). — Heterocera
Sumatrana 2 (6): 115–127)
Nässig, W.A., &
Holloway, J.D. (1989) A new species of Antheraea from Sumatra,
West Malaysia and Borneo (Lepidoptera, Saturniidae). — Heterocera
Sumatrana 2 (7): 185–190.
Paukstadt, L.H., & Paukstadt,
U. (1999): Die Beschreibung der Präimaginalstadien von Antheraea
(Antheraea) platessa W. Rothschild, 1903 von Nord Borneo [sic], sowie
Angaben zur Biologie und Ökologie (Lepidoptera: Saturniidae). — Galathea,
Ber. Krs. Nürnberger Entomol., Suppl. 6: 13–22.
Paukstadt, U., & Paukstadt,
L.H. (1999a): Anmerkungen zum taxonomischen Status von Actias selene
seitzi Kalis 1934 und
Actias selene vandenberghi Roepke 1956
(stat. rev.), sowie zur geographischen Verbreitung dieser Taxa in Südostasien
(Lepidoptera: Saturniidae). — Entomologische Zeitschrift 109 (12):
U., & Paukstadt, L.H. (1999b) [28. December]: Antheraea (Antheraea)
ulrichbroschi n. sp., eine neue Saturniide von West Malaysia [sic]
(Lepidoptera: Saturniidae). — Galathea, Ber. Krs. Nürnberger Entomol.,
Paukstadt, U., Paukstadt,
L.H., & Brosch, U. (1998):Taxonomische Änderungen und Anmerkungen
zu den Taxa der Gattung
Antheraea Hübner  von Sumatera
[sic], Indonesien (Lepidoptera: Saturniidae). — Entomologische Zeitschrift